Although a monophyletic group, male (XX/XY) and female heterogametic (WZ/ZZ) sex chromosome systems with a couple of variants like XX/X, Z/ZZ and multiple sex chromosome systems occur in insects. Molecular and morphological differences between X and Y or W and Z range from imperceptible to conspicuous. This article illustrates sex chromosome differentiation mainly in two fly species, Drosophila melanogaster and Megaselia scalaris, and in Lepidoptera. The earliest phases of XY evolution are present in the fly M. scalaris. Occasionally in this species, the male determining gene jumps to another chromosome, transforming the new host chromosome to a functional Y chromosome. Thus, in M. scalaris there are strains with virtually no XY differentiation (except for the sex determining function) and others with a moderate degree of differentiation. Base substitutions and alterations like sequence deletions, duplications, and insertions of mobile sequences mark the onset of molecular differentiation. Accumulation of molecular changes and coarser alterations are thought to lead to the morphological differences seen in WZ chromosome pairs of Lepidoptera. The W chromosome probably evolved in the most numerous clade of Lepidoptera, the Ditrysia, after it diverged from the common lepidopteran stem. Extant species display various degrees of molecular and morphological differentiation of the W chromosome, translocation or fusion with autosomes, and loss of the W.
Taste is important for food intake. The fetus first experiences taste through amniotic fluid, and later via mother’s milk. Early human experience with taste has a key importance for later acceptance of food. Dietary behavior is determined by the interaction of many different factors. The development of the olfactory and taste receptors begins at 7-8 weeks of gestation. An early sensitive period probably exists when flavor preference is established. Sweet taste is preferred in early childhood; this is the reason why children are at increased risk of over-consuming saccharides. Gustatory sensitivity declines with age. The threshold for the perception of each basic taste differs, and is established genetically. In this review, we summarize published data on taste preferences and its development and changes during life., Š. Podzimek, M. Dušková, Z. Broukal, B. Rácz, L. Stárka, J. Dušková., and Obsahuje bibliografii
The formation of ring species might provide an explanation of how speciation can occur despite ongoing gene flow. However, few species fit all of the criteria of a classic ring species that formed via isolation by distance around a barrier. Population genetic analyses and ecological niche models were used to examine a ring of song sparrow (Melospiza melodia) subspecies that surround the Sierra Nevada in North America. Eight models were compared that included both geography-based and ecology-based scenarios of ring formation. Song sparrows do fit some aspects of a classic ring species that formed via expansion around a barrier; however, admixture rather than complete reproductive isolation occurred when populations met at the terminus of the ring in southern California. Nichemodels show that variation among subspecies is likely to reflect adaptation to local conditions coupled with current limitations to gene flow across ecotones and that birds are likely to have expanded from a refugium in the southwestern United States. Given that simple isolation-based models often fail to explain many ring species patterns, alternative models that incorporate ecological factors might provide a better explanation of how most ring species formed. Isolation and subsequent partitioning of populations by ecotones can be important drivers of geographic variability in ring species.
In this paper, the feedback control for a class of bilinear control systems with a small parameter is proposed to guarantee the existence of limit cycle. We use the perturbation method of seeking in approximate solution as a finite Taylor expansion of the exact solution. This perturbation method is to exploit the "smallness" of the perturbation parameter ε to construct an approximate periodic solution. Furthermore, some simulation results are given to illustrate the existence of a limit cycle for this class of nonlinear control systems.
Effect algebras are very natural logical structures as carriers of probabilities and states. They were introduced for modeling of sets of propositions, properties, questions, or events with fuzziness, uncertainty or unsharpness. Nevertheless, there are effect algebras without any state, and questions about the existence (for non-modular) are still unanswered. We show that every Archimedean atomic lattice effect algebra with at most five blocks (maximal MV-subalgebras) has at least one state, which can be obtained by "State Smearing Theorem'' from a state on its sharp elements.
This paper deals with an experimental study of lubricant film in an experimental device (tribometer). Lubricating film is formed between a glass disc and steel ball on which modification for friction surfaces have been made. The experiment is aimed at both fully flooded EHL lubrication regime as well as the starved. This document also deals with a comparison of micro dents passage through a contact at different degrees of slip. and Obsahuje seznam literatury
This paper considers an exponential cost optimality problem for finite horizon semi-Markov decision processes (SMDPs). The objective is to calculate an optimal policy with minimal exponential costs over the full set of policies in a finite horizon. First, under the standard regular and compact-continuity conditions, we establish the optimality equation, prove that the value function is the unique solution of the optimality equation and the existence of an optimal policy by using the minimum nonnegative solution approach. Second, we establish a new value iteration algorithm to calculate both the value function and the ϵ-optimal policy. Finally, we give a computable machine maintenance system to illustrate the convergence of the algorithm.