Subject
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- evolution1
- feeding habits1
- female intrasexual fighting1
- female intrasexual mounting1
- habitat1
- habitat heterogeneity1
- historical factors1
- horned beetle1
- intraspecies competition1
- life history1
- male dimorphism1
- morphology1
- mouthparts1
- new combinations1
- nocturnal1
- particle feeding1
- paternal provisioning1
- phylogeny1
- post-copulatory processes1
- reproductive behaviour1